Characteristics of the Order (from Kramp 1961)

Scyphomedusae with umbrella margin cleft into lappets; without a coronal furrow and without pedalia; with (or without) hollow marginal tentacles and with marginal rhopalia; with a single central mouth opening provided with four large curtain-like or gelatinous lips; with gonads in sac-like folds of the endodermal wall of the subumbrella.